Model is underconstrained.
The following variables couldn't be defined:
 * V in component membrane
 * proton_motive_force in component membrane
 * pH_i in component membrane
 * pH_m in component membrane
 * JH_res in component proton_pump
 * JH_leak in component proton_leak
 * JH_F1 in component F0_F1_ATPase
 * J_ant in component ATP_ADP_exchange
 * J_uni in component Ca_uniporter
 * Jo in component proton_pump
 * A_res in component proton_pump
 * NAD_m in component proton_pump
 * NADH_m in component proton_pump
 * Jp_F1 in component F0_F1_ATPase
 * A_F1 in component F0_F1_ATPase
 * ATP_m in component F0_F1_ATPase
 * ADP_m_free in component F0_F1_ATPase
 * ATP_i in component ATP_ADP_exchange
 * ATP_m in component ATP_ADP_exchange
 * ADP_i in component ATP_ADP_exchange
 * ATP4_i in component ATP_ADP_exchange
 * ATP4_m in component ATP_ADP_exchange
 * ADP3_i in component ATP_ADP_exchange
 * ADP3_m in component ATP_ADP_exchange
 * Ca_i in component Ca_uniporter
 * J_Na_Ca in component Na_Ca_exchanger
 * d/dt V in component membrane
 * d/dt ADP_m in component ATP_ADP_exchange
 * d/dt Ca_m in component Na_Ca_exchanger